In 1915 France, Major Brand commands the 39th Squadron of the Royal Flying Corps. The young airmen go up in bullet-riddled "crates" and the casualty rate is appalling, but Brand can't make the "brass hats" at headquarters see reason. Insubordinate air ace Captain Courtney is another thorn in Brand's side...but finds the smile wiped from his face when he rises to command the squadron himself. Everyone keeps a stiff upper lip.

Following two fatal aircraft accidents aboard an Aircraft Carrier, a government agent is placed aboard the ship. His mission is to determine what may be at the root of these mishaps and prevent the inevitable third chain breaking accident which abruptly changes course when a sailor reports seeing one of the top ranking fighter pilots in a gay club. The aviator and three of his squadron mates and their spouses all become the subject of a deadly modern day witch hunt entangling them in a web of jealousy and deceit which tests their concept of love, friendship and honor and changes their lives forever.

During the Second World War, a special project is begun by the US Army Air Corps to integrate African American pilots into the Fighter Pilot Program. Known as the "Tuskegee Airman" for the name of the airbase at which they were trained, these men were forced to constantly endure harassement, prejudice, and much behind the scenes politics until at last they were able to prove themselves in combat.